Publicado 2026-07-23
Palabras clave
- Señales animales,
- Etología,
- Referencia funcional,
- Representación errónea
- Animals Signals,
- Ethology,
- Functional Reference,
- Misrepresentation

Esta obra está bajo una licencia internacional Creative Commons Atribución-NoComercial 4.0.
Resumen
Este artículo propone una explicación teleosemántica híbrida del significado en la comunicación entre animales (no humanos). Distingo dos familias de casos: señales sobre acontecimientos externos al emisor y señales sobre la identidad o la condición actual del emisor. Según el enfoque teleosemántico más influyente sobre la comunicación animal, tales señales representan las condiciones del mundo requeridas para que la conducta del receptor guiada por la señal resulte beneficiosa. Sostengo que, considerado por sí solo, este enfoque basado en el beneficio conduce a atribuciones de contenido poco plausibles, que se apartan tanto de la práctica etológica como de aquello que la propuesta se presenta como capaz de ofrecer en casos de comunicación animal. Para abordar este problema, recurro a un marco teleosemántico que desarrollé previamente para representaciones perceptivas simples, el cual aquí extiendo al caso de las señales animales. En particular, complemento el enfoque basado en el beneficio con una condición de producción de la señal que restringe el contenido a aquello que causalmente provoca la emisión de la señal por parte del emisor. El híbrido resultante asigna contenidos plausibles tanto para señales sobre acontecimientos del mundo externo como para señales sobre la identidad o la condición actual del emisor.
Citas
- Artiga, M. (2014). Signaling without cooperation. Biology & Philosophy, 29(3), 357-378. https://doi.org/10.1007/s10539-014-9436-0
- Artiga, M. (2021). Beyond black dots and nutritious things: A solution to the indeterminacy problem. Mind & Language, 36(3), 471-490. https://doi.org/10.1111/mila.12284
- Bradbury, J., & Vehrencamp, S. (1998). Principles of animal communication. Sinauer Associates.
- Caro, T. (2005). Antipredator defenses in birds and mammals. University of Chicago Press.
- Chaverri, G., & Gillam, E. (2010). Cooperative signaling behavior of roost location in a leaf-roosting bat. Communicative & Integrative Biology, 3(6), 599-601. https://doi.org/10.4161/cib.3.6.13277
- Cheney, D., & Seyfarth, R. (1990). How monkeys see the world: Inside the mind of another species. University of Chicago Press.
- Collett, T. S., & Land, M. F. (1978). How hoverflies compute interception courses. Journal of Comparative Physiology, 125, 191-204. https://doi.org/10.1007/BF00656597
- Cooley, J., Marshall, D., & Hill, K. (2018). A specialized fungal parasite (Massospora cicadina) hijacks the sexual signals of periodical cicadas (Hemiptera: Cicadidae: Magicicada). Scientific Reports, 8(1), 1432. https://doi.org/10.1038/s41598-018-19813-0
- Dretske, F. (1981). Knowledge and the flow of information. The MIT Press.
- Ewert, J.-P. (1980). Neuroethology: An introduction to the neurethological fundamentals of behavior. Springer. https://doi.org/10.1007/978-3-642-67500-3
- Ewert, J.-P., Burghagen, H., & Schürg-Pfeiffer, E. (1983). Neuroethological analysis of the innate releasing mechanism for prey-catching behavior in toads. In J. P. Ewert, R. R. Capranica & D. J. Ingle (Eds.), Advances in vertebrate neuroethology (pp. 413-475). Springer. https://doi.org/10.1007/978-1-4684-4412-4_21
- Fodor, J. (1990). A theory of content and other essays. The MIT Press. https://doi.org/10.7551/mitpress/6765.001.0001
- Godfrey-Smith, P. (1989). Misinformation. Canadian Journal of Philosophy, 19(4), 533-550. https://doi.org/10.1080/00455091.1989.10716781
- Ingle, D. (1976). Behavioral correlates of central visual function in anurans. In R. Llinás & W. Precht (Eds.), Frog neurobiology: A handbook (pp. 435-451). Springer. https://doi.org/10.1007/978-3-642-66316-1_13
- Johnsgard, P. (1965). Handbook of waterfowl behavior. Cornell University Press.
- Kruuk, H. (1964). Predators and anti-predator behaviour of the black-headed gull (Larus Ridibundus L.). Behaviour. Supplement, (11), III–129. http://www.jstor.org/stable/30039149
- Lettvin, J. Y., Maturana, H. R., McCulloch, W. S. & Pitts, W. H. (1959). What the frog’s eye tells the frog’s brain. Proceedings of the Institute of Radio Engineers, 47(11), 1940-1951. https://doi.org/10.1109/JRPROC.1959.287207
- Lloyd, J. (1965). Aggressive mimicry in Photuris: Firefly femmes fatales. Science, 149(3684), 653-654. https://doi.org/10.1126/science.149.3684.653
- Macedonia, J. M., & Evans, C. S. (1993). Essay on contemporary issues in ethology: Variation among mammalian alarm call systems and the problem of meaning in animal signals. Ethology, 93(3), 177-197. https://doi.org/10.1111/j.1439-0310.1993.tb00988.x
- Marler, P., Evans, C. S., & Hauser, M. D. (1992). Animal signals: Motivational, referential, or both? In H. Papoušek, U. Jürgens, & M. Papoušek (Eds.), Nonverbal vocal communication: Comparative and developmental approaches (pp. 66-86). Cambridge University Press.
- Martínez, M. (2013). Teleosemantics and indeterminacy. Dialectica, 67(4), 427-453. https://doi.org/10.1111/1746-8361.12039
- Millikan, R. G. (1984). Language, thought, and other biological categories. The MIT Press. https://doi.org/10.7551/mitpress/4124.001.0001
- Millikan, R. G. (1989). Biosemantics. The Journal of Philosophy, 86(6), 281-297. https://doi.org/10.2307/2027123
- Millikan, R. G. (1990a). Compare and contrast Dretske, Fodor, and Millikan on teleosemantics. Philosophical Topics, 18(2), 151-161. https://doi.org/10.5840/philtopics19901828
- Millikan, R. G. (1990b). Truth rules, hoverflies, and the Kripke-Wittgenstein paradox. The Philosophical Review, 99(3), 323-353. https://doi.org/10.2307/2185347
- Millikan, R. G. (1991). Speaking up for Darwin. In B. M. Loewer & G. Rey (Eds.), Meaning in mind: Fodor and his critics (pp. 151-164). Blackwell.
- Millikan, R. G. (2000). On clear and confused ideas: An essay about substance concepts. Cambridge University Press. https://doi.org/10.1017/CBO9780511613296
- Millikan, R. G. (2004). Varieties of meaning: The 2002 Jean Nicod lectures. The MIT Press. https://doi.org/10.7551/mitpress/7072.001.0001
- Millikan, R. G. (2017). Beyond concepts: Unicepts, language, and natural information. Oxford University Press. https://doi.org/10.1093/oso/9780198717195.001.0001
- Millikan, R. G. (2023). Teleosemantics and the frogs. Mind & Language, 39(1), 52-60. https://doi.org/10.1111/mila.12456
- Neander, K. (1991). Functions as selected effects: The conceptual analyst’s defense. Philosophy of Science, 58(2), 168-184. https://doi.org/10.1086/289610
- Neander, K. (1995). Misrepresenting & malfunctioning. Philosophical Studies, 79(2), 109-141. https://doi.org/10.1007/BF00989706
- Neander, K. (2006). Content for cognitive science. In G. Macdonald & D. Papineau (Eds.), Teleosemantics: New philosophical essays (pp. 167-194). Oxford University Press. https://doi.org/10.1093/oso/9780199270262.003.0009
- Neander, K. (2017). A mark of the mental: In defense of informational teleosemantics. The MIT Press. https://doi.org/10.7551/mitpress/7387.001.0001
- Papineau, D. (1984). Representation and explanation. Philosophy of Science, 51(4), 550-572. https://doi.org/10.1086/289205
- Papineau, D. (1993). Philosophical naturalism. Blackwell.
- Pietroski, P. (1992). Intentionality and teleological error. Pacific Philosophical Quarterly, 73(3), 267-282. https://doi.org/10.1111/j.1468-0114.1992.tb00339.x
- Price, C. (2001). Functions in mind: A theory of intentional content. Oxford University Press. https://doi.org/10.1093/acprof:oso/9780199242009.001.0001
- Rogers, L. J., Stewart, L., & Kaplan, G. (2018). Food calls in common marmosets, Callithrix jacchus, and evidence that one is functionally referential. Animals, 8(7), 99. https://doi.org/10.3390/ani8070099
- Schulte, P. (2012). How frogs see the world: Putting Millikan’s teleosemantics to the test. Philosophia, 40, 483-496. https://doi.org/10.1007/s11406-011-9358-x
- Schulte, P. (2018). Perceiving the world outside: How to solve the distality problem for informational teleosemantics. The Philosophical Quarterly, 68(271), 349-369. https://doi.org/10.1093/pq/pqx052
- Shea, N. (2007). Consumers need information: Supplementing teleosemantics with an input condition. Philosophy and Phenomenological Research, 75(2), 404-435. https://doi.org/10.1111/j.1933-1592.2007.00082.x
- Shea, N. (2018). Representation in cognitive science. Oxford University Press. https://doi.org/10.1093/oso/9780198812883.001.0001
- Seyfarth, R. M., Cheney, D. L., & Marler, P. (1980a). Vervet monkey alarm calls: Semantic communication in a free-ranging primate. Animal Behaviour, 28(4), 1070-1094. https://doi.org/10.1016/S0003-3472(80)80097-2
- Seyfarth, R. M., Cheney, D. L., & Marler, P. (1980b). Monkey responses to three different alarm calls: Evidence of predator classification and semantic communication. Science, 210(4471), 801-803. https://doi.org/10.1126/science.7433999
- Smith, J. M., & Harper, D. (1995). Animal signals: Models and terminology. Journal of Theoretical Biology, 177(3), 305-311. https://doi.org/10.1006/jtbi.1995.0248
- Smith, J. M., & Harper, D. (2003). Animal signals. Oxford University Press. https://doi.org/10.1093/oso/9780198526841.001.0001
- Tinbergen, N. (1948). Social releasers and the experimental method required for their study. The Wilson Bulletin, 60(1), 6-47.
- Withrington, E. (2024). Frogs recognize prey: A causal-behavioral teleosemantics. Biology & Philosophy, 39(6), article 35. https://doi.org/10.1007/s10539-024-09967-9
